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Angomonas deanei
''Angomonas deanei'' is a flagellated trypanosomatid protozoan. It is an obligate parasite in the gastrointestinal tract of insects, and is in turn a host to symbiotic bacteria. In fact the endosymbiont maintains a permanent mutualistic relationship with the bacterium such that the bacterium is no longer able to reproduce and survive on its own.
The species name was accepted as ''Crithidia deanei'' until 2011, when phylogenetic analysis revealed that it belongs to the genus ''Angomonas'', thereby becoming ''Angomonas deanei''. The symbiotic bacterium is a member of the β-proteobacterium that descended from the common ancestor with the genus ''Bordetella'', or more likely, ''Taylorella''. The two organisms have depended on each other so much that the bacterium cannot reproduce and the protozoan can no longer infect insects when they are isolated. Thus, this organismal association is a fine model for the evidence of the endosymbiotic theory in nature, which explains the origin of eukaryotic cell organelles such as mitochondria and plastids from individual prokaryotes.
==Structure==
The body of ''Angomonas deanei'' is elliptical in shape, with a prominent tail-like flagellum at its posterior end for locomotion. The bacterial endosymbiont is inside its body and is surrounded by two membranes typical of Gram-negative bacteria, but its cell membrane presents unusual features, such as the presence of phosphatidylcholine, a major membrane lipid (atypical of bacterial membranes), and the highly reduced peptidoglycan layer, which shows reduced or absence of rigid cell wall. The cell membrane of the protozoan host contains an 18-domain β-barrel porin, which is a characteristic protein of Gram-negative bacteria. In addition it contains cardiolipin and phosphatidylcholine as the major phospholipids, while sterols are absent. Cardiolipin is a typical lipid of bacterial membranes; phosphatidylcholine, on the other hand, is mostly present in symbiotic prokaryotes of eukaryotic cells. For symbiotic adaptation, the host trypanosome has undergone alterations such as reduced paraflagellar rod, which is required full motility of the bacterial flagella. Yet the paraflagellar rod gene ''PFR1'' is fully functional. The bacteria are known to provide essential nutrients to the host, and provide electron transport system for the production of cellular energy, the ATP molecules through its glycosomes.〔 In return the protozoan offers its enzymes for the complete metabolic pathways for the biosysnthesis of amino acids, lipids and nucleotides, that are absent in the protozoan. Thus the two organisms intimately share and exchange their metabolic systems. When the protozoan is removed off of the bacterium using antibiotics, it loses its infectivity to insects,〔 obviously due to the altered glycosylphosphatidylinositol (gp63) in the protozoan flagellum.
The cellular reproduction shows a strong synergistic adaptation between the bacterium and the protozoan. As each endosymbiont is each of a single bacterium and a protozoan, and each daughter cell contains the same number, the two cells divide in a coordinated process. The bacterium divides first, followed by the protozoan organelles, and lastly the nucleus.
抄文引用元・出典: フリー百科事典『 ウィキペディア（Wikipedia）』
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